main culprits

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1.
The main culprits were vitamin A, beta carotene and vitamin E; vitamin C and selenium seemed to have no meaningful effect.
罪魁祸首就是维生素A、β-胡萝卜素和维E；维C和硒也没显示有明显效果。
article.yeeyan.org
2.
But urban runoff and septic seepage are not always the main culprits behind microbial pollution.
不过，都市迳流及化粪池的渗漏，并非一直是微生物污染背后的主要凶手。
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3.
Eye makeup can be one of the main culprits of styes, which can take months to get rid of.
眼妆可能是导致眼病的罪魁祸首，麦粒肿（眼病）一般要几个月时间才能痊愈。
article.yeeyan.org
4.
The agencies are neither the only, nor indeed the main, culprits for the subprime crisis.
确实，信评机构不是造成信贷危机的罪魁祸首。
blog.sina.com.cn
5.
The twin ports of Los Angeles and Long Beach are among the main culprits for generating pollution in the area.
加州洛杉矶和长堤的海港是这一地区空气污染的罪魁祸首之一。
www.putclub.com
6.
Undoubtedly, as legal guardian and early mentor, parents cannot evade the responsibility and are the main culprits.
毫无疑问，作为合法的监护人和早期的指导者，父母是主要的罪魁祸首，无法逃避责任。
blog.sina.com.cn
7.
Swordfish, shark, tilefish, fresh tuna steak and king mackerel are the main culprits.
箭鱼、鲨鱼、方头鱼、新鲜的金枪鱼排和鲭鱼等。
article.yeeyan.org
8.
It is these collaborators the government wants to try, not the main culprits in the former West Pakistan army.
孟加拉国政府想要审判的就是这些通敌分子，儿不是西巴基斯坦军队里的重要战争罪犯。
www.ecocn.org
9.
There are three main culprits:
罪魁祸首主要有三个：
article.yeeyan.org